Red grouper excavate holes in the limestone sea floor filled in by sediment, providing habitats for corals, sponges, lobster, shrimp, snapper, and other species, according to a new study led by Felicia Coleman (2000) and co-authored by Selena Heppell (2006). Susan Williams (2000), who collaborated with Coleman previously, called the groupers "'Frank Lloyd Wrights' of the sea floor", saying that "if the groupers are overfished, the suite of species that depends on them is likely to suffer."